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Prof. Itay Mayrose Lab – Plant Evolution, Bioinformatics, & Comparative Genomics







CCDB is a community resource. If you are aware of any missing data or if any of the counts are erroneous, please submit new counts, or report erroneous counts.


CCDBAngiospermsCannabaceaeHumulusHumulus lupulus L.

52 chromosome counts in Humulus lupulus L.:


Name Accepted Name Gametophytic(n) Sporophytic(2n) Data Source reference
  Humulus lupulus L. Humulus lupulus L.   /20/ Slovakia Database Májovský J. et al., 1970b: Index of chromosome numbers of Slovakian flora Part 2. - Acta Fac. Rerum Nat. Univ. Comen., Bot. 18: 45-60.
  Humulus lupulus L. Humulus lupulus L.   20 IAPT/IOPB IAPT/IOPB Chromosome Data 8
!   Humulus lupulus L. Humulus lupulus L.   20 British Isles cytology Dempsey, R.E., Gornall, R.J., Bailey, J.P. Watsonia 20: 63-66 (1994)
!   Humulus lupulus L. Humulus lupulus L.   20 flora of NW Europe http://wbd.etibioinformatics.nl/bis/flora.php?selected=beschrijving&menuentry=soorten&id=1740
!   Humulus lupulus L. Humulus lupulus L.   20 Flora Europaea Polya, 1949
  Humulus lupulus L. Humulus lupulus L.   20 Poland Database Pogan E., Czapik R., Jankun A., Kuta E. et al. 1982 a. Further studies in chromosome numbers of Polish Angiosperms. Part XV. Acta Biol. Cracov. Ser. Bot. 24, 91-126.
  Humulus lupulus L. Humulus lupulus L.   20 Poland Database Grabowska-Joachimiak A., Sliwinska E., Pigu?a M., Skomra U., Joachimiak A.J. 2006. Genome size in Humulus lupulus L. and H. japonicus Siebold & Zucc. (Cannabaceae). Acta Soc. Bot. Pol. 75, 207-214.
  Humulus lupulus L. Humulus lupulus L.   30 Poland Database Grabowska-Joachimiak A., Sliwinska E., Pigu?a M., Skomra U., Joachimiak A.J. 2006. Genome size in Humulus lupulus L. and H. japonicus Siebold & Zucc. (Cannabaceae). Acta Soc. Bot. Pol. 75, 207-214.
  Humulus cordifolius Miq. Humulus lupulus L.   20 IPCN online Probatova, N. S. & A. P. Sokolovskaya. 1989. Chromosome numbers in vascular plants from Primorye Territory, the Amur region, Sakhalin, Kamchatka and the Kuril Islands. Bot. Žhurn. (Moscow & Leningrad) 74: 120–123.
  Humulus lupulus L. Humulus lupulus L.   20 IPCN online Pogan, E., R. CZAPIK, A. Jankun & E. Kuta. 1982. Further studies in chromosome numbers of Polish angiosperms. Part XV. Acta Biol. Cracov., Ser. Bot. 24: 113–126.
  Humulus lupulus L. Humulus lupulus L.   20 IPCN online Dempsey, R. E., R. J. Gornall & J. P. Bailey. 1994. Contributions to a cytological catalogue of the British and Irish flora, 4. Watsonia 20: 63–66.
  Humulus lupulus L. Humulus lupulus L.   20 IPCN online Druskovic, B. & M. Lovka. 1995. IOPB chromosome data 9. Int. Organ. Pl. Biosyst. Newslett. (Zurich) 24: 15–19.
  Humulus lupulus L. Humulus lupulus L.   20 IPCN online Hill, L. M. 1995. IOPB chromosome data 10. Int. Organ. Pl. Biosyst. Newslett. (Zurich) 25: 8–9.
  Humulus lupulus L. Humulus lupulus L.   20, 40 IPCN online Stepanov, N. V. & E. N. Muratova. 1995. Chromosome numbers of some taxa of higher plants of Krasnoyarsk territory. Bot. Žhurn. (Moscow & Leningrad) 80(6): 114–116.
  Humulus lupulus L. Humulus lupulus L.   20 IPCN online Dobea, C., B. Hahn & W. Morawetz. 1997. Chromosomenzahlen zur Gefässpflanzen-Flora Österreichs. Linzer Biol. Beitr. 29(1): 5–43.
  Humulus lupulus L. Humulus lupulus L.   20 IPCN online Jav|0urková-Jarolímová, V. 1992. In J. M|3esí|3ccek & V. Jav@0urková-Jarolímová, List of Chromosome Numbers of the Czech Vascular Plants. Academia, Praha.
  Humulus lupulus L. Humulus lupulus L.   20, 21 IPCN online Shephard, H. L., J. S. Parker, P. Darby & C. C. Ainsworth. 2000. Sexual development and sex chromosomes in hop. New Phytol. 148: 397–411.
  Humulus lupulus L. Humulus lupulus L.   20 IPCN online Stoian, V., N. Constantin & A. Cuniţă. 1998. Sex chromosomes in some flowering plants of the Romanian flora. Rev. Roumaine Biol., Sér. Biol. Vég. 43(1): 11–19.
  Humulus lupulus L. Humulus lupulus L.   40 IPCN67-71 HAUNOLD, A. 1968. Venetian turpentine as an aid in squashing and concomitant production of durable chromosome mounts. stain Technol. 43: 153-156.
  Humulus lupulus L. Humulus lupulus L.   20, 40 IPCN67-71 MAJOVSKY, J., ET. AL. 1970. Index of chromosome numbers of Slovakian flora. (Part 2). Acta Fae. Rerum Nat. Univ. Comenianae Bot. 18: 45-60.
  Humulus lupulus L. Humulus lupulus L.   20 IPCN75-78 lfotegi 1976
!   Humulus lupulus L. Humulus lupulus L.   20 Kumar, 1987 Tournois, J. 1914 ; Winge,(>. 1914. 1917a, b, 1923, 1929 ; Bartlett, H.H. 1915 ; Kihara, H. 1929 ; Sinoto, Y. 1929a, b ; Kuhn (Tischler, G. 1931) ; Tischler, G. 1934; Nakajima, G. 1937b ; Ono, T. 1937, 1959 ; Golu- binskii, E.N. (Fedorov); P6lya, L. 1949 ; Hara, H. 1952 ; USve, A. 1954b ; Jacobsen, P. 1957 ; Neve, R.A. 1958
!   Humulus lupulus L. Humulus lupulus L.   20, 40 Kumar, 1987 Majovsky, J. et al. ~970b
!   Humulus lupulus L. Humulus lupulus L.   40 Kumar, 1987 Haunold, A. 1968
!   Humulus lupulus L. Humulus lupulus L.   20 eFlora  
  Humulus lupulus L. Humulus lupulus L.   20 Fedorov, 1974 Tournois 1914
  Humulus lupulus L. Humulus lupulus L.   20 Fedorov, 1974 T. 1921-f.922)
  Humulus lupulus L. Humulus lupulus L.   20 Fedorov, 1974 Winge 1914, 1917a, b, 1923, 1929
  Humulus lupulus L. Humulus lupulus L.   20 Fedorov, 1974 Bartlett 1915c
  Humulus lupulus L. Humulus lupulus L.   20 Fedorov, 1974 Kihara 1929
  Humulus lupulus L. Humulus lupulus L.   20 Fedorov, 1974 Sinot& 1929a, b
  Humulus lupulus L. Humulus lupulus L.   20 Fedorov, 1974 Kuhn (T. 1931)
  Humulus lupulus L. Humulus lupulus L.   20 Fedorov, 1974 Tischler 1934
  Humulus lupulus L. Humulus lupulus L.   20 Fedorov, 1974 Nakajima 1937b
  Humulus lupulus L. Humulus lupulus L.   20 Fedorov, 1974 Ono T. 1937 roeyOBBCKBi 1947
  Humulus lupulus L. Humulus lupulus L.   20 Fedorov, 1974 P6lya 1949
  Humulus lupulus L. Humulus lupulus L.   20 Fedorov, 1974 Hara H. 1952
  Humulus lupulus L. Humulus lupulus L.   20 Fedorov, 1974 Liive A. 1954b 28*
!   Humulus lupulus L. Humulus lupulus L.   20 Fedorov, 1974 Jacobsen 1957
!   Humulus lupulus L. Humulus lupulus L.   20 Fedorov, 1974 Neve 1958
!   Humulus lupulus L. Humulus lupulus L.   2n = 20 Austria_checklist Dobe?, Ch., Hahn, B., & Morawetz, W. 1997. Chromosomenzahlen zur Gef??pflanzen-Flora ?sterreichs. Linzer Biol.Beitr. 29[1], 5-43.
!   Humulus lupulus L. Humulus lupulus L. 10 20 Cave1957 Jacobsen
  Humulus lupulus L. Humulus lupulus L.   20 CromoCat 2015 Cheshmedziev, I. V. (1994). Reports (313-366). Fl. Medit. 4: 269-279.
  Humulus lupulus L. Humulus lupulus L.   20 CromoCat 2015 Dobeš, C., Hahn, B. & Morawetz, W. (1997). Chromosomenzahlen zur Gefässpflanzen-Flora Österreichs. Linzer Biol. Beitr. 29(1): 5-43.
  Humulus lupulus L. Humulus lupulus L.   20 CromoCat 2015 Druskovic, B. & Lovka, M. (1995). IOPB Chromosome Data 9 (ed. C.A. Stace). Int. Organ. Pl. Biosyst. Newslett. 24: 15-19.
  Humulus lupulus L. Humulus lupulus L.   20 CromoCat 2015 Gabrielan, E. & Zohary, D. (2004). Wild relatives of food crops native to Armenian and Nakhichevan Fl. Medit. 14: 5-80
  Humulus lupulus L. Humulus lupulus L.   20 CromoCat 2015 Hill, L. M. (1995a). IOPB Chromosome Data 10 (ed. C.A. Stace). Int. Organ. Pl. Biosyst. Newslett. 25: 8-9.
  Humulus lupulus L. Humulus lupulus L.   20 CromoCat 2015 M?sí?ek J. et Jarolímová-Jav?rková V. (1992). List of chromosome numbers of the Czech vascular plants. Academia, Praha. p. 1-144.
  Humulus lupulus L. Humulus lupulus L.   20 CromoCat 2015 Probatova, N.S., Seledets, V.P., Rudyka, E.G., Gnutikov, A.A., Kozhevnikova, Z.V. & Barkalov V.Y. (2009). IAPT/IOPB Chromosome Data 8 Taxon 58 (4): 1284-1288
  Humulus lupulus L. Humulus lupulus L.   21 CromoCat 2015 Shephard, H. L., J. S. Parker, P. Darby & C. C. Ainsworth. (2000). Sexual development and sex chromosomes in hop New Phytologist 148: 397-411
  Humulus lupulus L. Humulus lupulus L.   40 CromoCat 2015 Stepanov, N. V. & E. N. Muratova. (1995). Chromosome numbers of some taxa of higher plants of Krasnoyarsk territory. Bot. Žurn. 80(6): 114-116.
!   Humulus lupulus L. Humulus lupulus L.   20 CromoCat 2015 Löve, A. & Löve, D. (1982a). Reports [In Löve, A. (ed.), IOPB chromosome numbers reports LXXIV]. Taxon 31(1): 124-126.